Invasion History
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First Non-native North American Tidal Record: 1924First Non-native West Coast Tidal Record: 1924
First Non-native East/Gulf Coast Tidal Record:
General Invasion History:
Ocenebra inornata is native to the Northwest Pacific, from the southern Kurile Islands in the Sea of Okhotsk through the Sea of Japan and the Yellow Sea, south to Jindo Island, South Korea (Golikov et al. 1976; Huang et al. 2001, Kil et al. 2005). It has been introduced to the Pacific Coast of North America (California to British Columbia) and the Atlantic Coast of Europe (France to Denmark). It has been transported around the world with Pacific Oyster (Crassostrea gigas) transplants.
North American Invasion History:
Invasion History on the West Coast:
Ocenebra inornata was first collected on the West coast in 1924 in Samish Bay, Washington (WA) on newly transplanted Pacific Oysters (Crassostrea gigas) from Japan (Kincaid 1947, cited by Carlton 1979). It was subsequently found in many nearby areas where oysters were cultured, including Boundary Bay, British Columbia (BC) in 1931 (Sherwood 1931, cited by Carlton 1979), Puget Sound in 1932 (Galtsoff 1932, cited by Carlton 1979; Cohen et al. 1998), and Ladysmith, Vancouver Island, BC in 1934 (Elsey 1934, cited by Carlton 1979). In 1939, they appeared in Willapa Bay, WA (Hanna 1966, cited by Carlton 1979), where they are still abundant (Buhle and Ruesink 2009). They were found further south in Netarts Bay, Oregon in 1934 (Squire 1972, cited by Carlton 1979; Carlton 1992; Martel et al. 2004b), and in Tomales Bay, California (CA) in 1941 (Copeland 1941, cited by Carlton 1979), where its presence is reported to be variable (McLean, in Carlton 2007). In 1966, two live specimens of O. inornata were collected in Morro Bay, CA (Carlton 1979), and in 2007, several live specimens were found in Humboldt Bay, CA (Mello, personal communication; Eberle, personal communication; Carlton, personal communication). This snail has direct development in egg capsules, and thus tends to have a localized distribution around oyster-growing areas (Garcia-Meunier et al. 2002).
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Invasion History Elsewhere in the World:
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In the Northeast Atlantic, Ocenebra inornata was discovered in the Baie de Marennes-Oléron (Bay of Biscay), France in 1995 (Garcia-Meunier et al. 2002). Based on molecular analyses, O. inornata was probably introduced to France with imports of Pacific Oysters from Washington State, with later genetic contributions of oysters imported directly from Japan (Martel et al. 2004a). By 2002, it had spread northward to at least nine other oyster-culture areas on the Atlantic Coast of France, as far as Quiberon in south Brittany (Martel et al. 2004b). Ocenebra inornata was collected in Jutland, Denmark in 2006 (Lützen et al. 2011); Yerseke in the Oosterschelde estuary, Netherlands in 2007; and Tuttelhoek, Netherlands in 2008 (Faasse and Ligthart 2009). In the Netherlands, it was initially misidentified as O. erinacea (Faasse and Ligthart 2009).